Nepenthes hemsleyana
Synonymy
Nepenthes hemsleyana in Engler, Pflanzenr. IV. 111 (Heft 36): 61. 1908 [The specific epithet honours Kew botanist William Botting Hemsley (1843 –1924), who described N. macfarlanei Hemsl. in 1905.D]. sec. Scharmann & Grafe 2013
- Lectotype (designated by Scharmann, M. & Grafe, T. U. 20131): Borneo, Sarawak, Lawas River, 1877, F.W.T. Burbidge s.n. (K: K00065148) http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000651485
- =Nepenthes rafflesiana var. subglandulosa in Int. J. Bot. 2(4): 348 (-349; fig. 23). 2006 syn. sec. Scharmann & Grafe 2013
- –Nepenthes rafflesiana var. elongata in Kew Bull. 1897: 405. 1897, nom. nud., syn. sec. Clarke & al. 20112
Description
Terrestrial climber to 6 m tall. Stems terete, up to 8 mm diam. Internodes 0.5 –1 cm on rosettes, 10 –15 cm on climbing stems. Leaves of rosettes chartaceous to thin-coriaceous, petiolate, the petioles narrow, lacking wings and canaliculate, up to 12 cm long, sheathing the stem for up to 1/2 – 3/4 of its circumfer- ence, not decurrent. Leaf blades oblong, up to 30 cm long, base abruptly contracted into the petiole; apex obtuse, acuminate, tendril insertion simple. Longitudinal nerves 3 – 5 on each side of the midrib, pennate nerves spreading towards the margins, but often inconspicuous. Tendrils uncoiled, up to 35 cm long. Leaves of climbing stems similar to those of the rosettes but smaller; petioles up to 10 cm long, not winged, sheathing the stem and not decurrent; leaf blades arising gradually from the petiole, oblong, up to 18 cm long, apex acuminate, tendril insertion simple. Lower pitchers up to 20 cm high, up to 5 cm wide, thin-chartaceous, arising abruptly from the tendril, broadly ovoid in the lower 1/3 with a pronounced hip, cylindrical above, nar- rowing slightly towards the mouth. Inner surfaces of the ovoid portion below the hip glandular throughout, surfaces above the hip covered in a layer of wax crystals. Two fringed wings, up to 3 cm wide (widest at the base), bearing multicellular fringe elements up to 12 mm long, run from the bottom of the pitcher to the mouth at the front. Mouth round, oblique, concave, rising at the rear into a distinct neck. Peristome sub-cylindrical, up to 8 mm wide at the front and sides, up to 12 mm wide near the apex. Outer surface entire, inner surface with distinct teeth up to 5 mm long; ribs up to 1 mm apart, up to 0.5 mm wide. Lid broadly ovate, base cordate, up to 6 cm long, up to 5 cm wide, lacking appendages on the lower surface. Large, crater-like nectar glands, up to 0.5 mm wide, scattered sparsely to densely around the outer lower surfaces. Spur simple, up to 10 mm long. Upper pitchers 18 – 25 cm high, 3 – 5 cm wide, thin-chartaceous, arising very gradually from the hanging end of the tendril, narrowly infundibular in the lower 1/3, becoming noticeably broader towards the hip, which is located 1/2 – 2/3 of the way up the pitcher; cylindrical above the hip to the peristome. Mouth and peristome similar to the lower pitchers, the latter sometimes with a slight kink at the front. Glandular region covers the entire inner surface below the hip; cylindrical portion above the hip covered with wax crystals. Lid ovate, generally not cordate at the base, up to 6 cm long by 4 cm wide, no appendages on the lower surface. Large, crater-like nectar glands, up to 0.5 mm wide, scattered sparsely around the outer lower surfaces. Spur simple, up to 10 mm long. Male inflorescence a raceme, peduncle up to 12 cm, rachis up to 30 cm, partial peduncles 1-flowered, bracts usually absent, pedicels 12 –15 mm long, tepals elliptic, up to 7 by 5 mm; androphore 5 – 6 mm long, anther head 1.2 by 2 mm. Female inflorescence similar in structure to the male, peduncle up to 12 cm, rachis up to 20 cm, partial peduncles 1-flowered, lacking bracts. Valves of fruits 50 by 10 mm. Indumentum of stem, midribs, lower surface of the leaf blade and inflorescences from base of peduncle to lower surface of the tepals white or grey arachnoid; lower surface of leaf and outer surface of pitchers with minute grey stellate hairs. Other surfaces glabrous. Colour of the leaves and pitchers drying to light green or straw brown, stem white, flowers dark brown. Pitchers on living plants pure light green throughout, or green with red-purple specks on the outer surfaces and lid; peristome green throughout to striped with varying degrees of red and green. Stems whitish grey, leaves dull green.A
Distribution (General)
Borneo: Baram district and Bintulu area of Sarawak, and Belait and Tutong districts of BruneiB
Habitat
Terrestrial climber in peat swamp forest, kerangas and kerapah habitats, occasionally locally abundant in disturbed kerangas or along kerangas forest ecotones; below 200 metres a.s.l.C
Bibliography
A. Clarke, C. M., Moran, J.A. & Lee, C.C. 2011: Nepenthes baramensis (Nepenthaceae) – a new species from north-western Borneo. – Blumea 56(3): 229-233 (as Nepenthes baramensis in Blumea 56(3): 230 (229-233; fig. 1). 2011)
B. Scharmann, M. & Grafe, T. U. 2013: Reinstatement of Nepenthes hemsleyana (Nepenthaceae), an endemic pitcher plant from Borneo, with a discussion of associated Nepenthes taxa. – Blumea 58(1): 8-12
C. Clarke, C. M., Moran, J.A. & Lee, C.C. 2011: Nepenthes baramensis (Nepenthaceae) – a new species from north-western Borneo. – Blumea 56(3): 229-233 (as Nepenthes baramensis in Blumea 56(3): 230 (229-233; fig. 1). 2011)
Specimens
| Country | Date | Collector + collecting number | Herbaria | Type | Scan | Derivatives | |
|---|---|---|---|---|---|---|---|
| 1877 | Burbidge, F. W. T. s.n. | K |  | ||||
| Citation: Borneo, Sarawak, Lawas River, 1877, F.W.T. Burbidge s.n. (K K00065148) Specimen summary: K K00065148 Preferred stable URI: http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000651485 Lectotype of Nepenthes hemsleyana Macfarl. | |||||||