Nepenthes ×hookeriana

Nepenthes ×hookeriana

Synonymy

Nepenthes ×hookeriana H.Low ex Mast. in Gard. Chron. n.s. 16: 812 (& 813, fig. 157). 1881 sec. Cheek & Jebb 2001
  • Nepenthes hookeriana H.Low ex Lindl. in Gard. Chron. 1848: 87. 1848, nom. nud., syn. sec. Berendsohn 2017+3
  • Nepenthes hookeriana H.Low ex W.H.Baxter, Suppl. Hort. Brit.: 593. 1850, nom. nud., syn. sec. Berendsohn 2017+
  • Nepenthes loddigesii W.H.Baxter, Suppl. Hort. Brit.: 593. 1850, nom. nud., syn. sec. Berendsohn 2017+4
  • Nepenthes hookeri Alphand, Prom. de Paris: ic. s.n. 1867–1873, nom. nud., syn. sec. Berendsohn 2017+
  • Nepenthes hookeri Alphand ex Hook.f. in de Candolle, Prodr. 17: 96. 1873, , nom. inval., syn. sec. Berendsohn 2017+5,6
  • 3. Type: Low (not located) Borneo, Sarawak, 4. Type: not located, 5. in syn. Nepenthes rafflesiana, 6. Type: Alphand, Prom. de Paris, cum ic. (n.v.).
  • =? Nepenthes ×excelsior B.S.Williams in Garden Lond. 28: 463. 1885 syn. sec. Cheek & Jebb 2001
  • Nepenthes rafflesiana var. excelsior (Lindl.) Beck in Wiener Ill. Gart.-Zeitung 20: 147. 1895 syn. sec. Cheek & Jebb 2001
  • =Nepenthes rafflesiana var. hookeriana (Lindl.) Beck in Wiener Ill. Gart.-Zeitung 20: 147. 1895 syn. sec. Cheek & Jebb 2001
  • Nepenthes hookeriana H.Low, Sarawak: 68. 1848, nom. nud., syn. sec. Cheek & Jebb 2001

Other sources

Nepenthes hookeriana H.Low ex Lindl.1: Clarke, C.M. 1997: Nepenthes of Borneo: 153, f. 106 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes loddigesii W.H.Baxter2: Beck von Mannagetta und Lerchenau, G. 1895: – Wiener Ill. Gart.-Zeitung 20: 227 (as Nepenthes loddigesii W.H.Baxter2)
Nepenthes hookeriana H.Low ex Lindl.1: Lindley, J. 1892: – Gard. Chron.: 561, ic. 557 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Danser, B.H. 1928: – Bull. Jard. Bot. Buitenzorg, sér. 3, 9: 309, f. 9 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Jebb, M.H.P. & Cheek, M.R. 1997: A skeletal revision of Nepenthes. – Blumea 42(1): 1-106: 50 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Burbidge, F.W.T. 1904: – Flora & Sylva 2: 111 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Nicholson, G. 1887: Ill. Dict. Gard. 4: 436 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Macfarlane, J.M. 1908: pp. 1-92. – In: Engler, A., Pflanzenreich IV. 111 (Heft 36): 34 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Shivas, R.G. 1984: Pitcher Plants of Peninsula Malaysia & Singapore: 33 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Phillipps, A. & Lamb, A.L. 1996: Pitcher Plants of Borneo: 94, f. 7 (as Nepenthes hookeriana H.Low ex Lindl.1)
Nepenthes hookeriana H.Low ex Lindl.1: Masters, M.T. 1881: – Gard. Chron. n.s.: 812, f. 157 (as Nepenthes hookeriana H.Low ex Lindl.1)

Description

In morphology Nepenthes ×hookeriana is intermediate between the parental species. The leaf blade exhibits the typical venation of N. ampullaria, with the longitudinal nerves in the outer 1/2 of the blade only, and a shortly petiolate base; the lower pitchers are urceolate with broad pitcher wings and a broad, rounded peristome, but this is not developed into the long apical neck of N. rafflesiana; the lid is oblong to oblong-ovate, with a blunt or notched apex, and two prominent lateral veins, the lid glands are distributed throughout, unlike those of N. rafflesiana, which are densest near the margins and absent from the centre.A

Notes

Nepenthes x hookeriana Lindl. in the Gardeners’ Chronicle is merely a name in a list of species, referring to the name in Low’s book. Hugh Low, however, accidentally, or otherwise, had described what we know as N. rafflesiana as Nepenthes x hookeriana and vice versa in his book Sarawak, its Inhabitants and Productions (1848). Masters was the first author to note this in the Gardeners’ Chronicle II (1881) 818, f. 157, where he gives the first full description and illustration of Nepenthes x hookeriana. However the species still effectively remained dubious taxonomically (even though its facies were well understood in horticultural circles), until Macfarlane’s revision. Macfarlane (op. cit. 1908) cites several specimens, among them a Low collection from Sarawak, which would seem the most appropriate choice for a lectotype, but we have not been able to locate this specimen. Nepenthes loddigesii is included on the authority of Macfarlane, but no type material has been located.
Along with another naturally occurring hybrid, Nepenthes ×trichocarpa, this taxon is widespread, albeit scarce. The numbers of plants present in a given population are often small and they tend to be very localised. It is possible that hybrids can only survive in marginal or disturbed habitats, since the ecologies of the two parental species is not identical. Other hybrids such as Nepenthes ×kinabaluensis and Nepenthes ×trusmadiensis are highly restricted in their distribution, and their identification is not problematic on a Malesian scale.A
Some notes on nomenclature of Nepenthes ×hookeriana:
- Cheek & Jebb (2001) cite this hybrid with Lindley as the author, but see their note.
- H.Low in Sarawak: 68, 1848 publishes N. hookeriana as a new species, different from N. rafflesiana, but mixes up the two descriptions.
- Nepenthes hookeriana H.Low ex Lindl. in Gard. Chron. 1848: 87
is referring the name to Low but it does not cite N. rafflesiana and thus could be interpreted as following Low’s erroneous description. So, since there is no direct reference to an earlier description, this is considered a nomen nudum.
- Nepenthes hookeriana H.Low ex W.H.Baxter, Suppl. Hort. Brit.: 593. 1850 is again referring the name to Low and cites N. rafflesiana in the same table. Still, as Low’s error is not pointed out, this is another nomen nudum.
- Nepenthes loddigesii W.H.Baxter, Suppl. Hort. Brit.: 593. 1850 is another entry in the same table, without any diagnosis, not referring to an earlier description; another nomen nudum.
- Nepenthes hookerii Alphand, Prom. de Paris: ic. s.n. 1868.
(The last plate in section "Flore Ornamental, planches", see
http://gallica.bnf.fr/ark:/12148/bpt6k6276852z/f483.item.r=Nepenthes.langFR.zoom). Plate with name, no further data.
- Nepenthes hookeri Alphand ex Hook.f. in A.DC., Prodr. 17: 96. 1873 cites the former in synonymy – this does not validate the name. B

Distribution (General)

Sumatra, Peninsular Malaysia, Singapore, Borneo.A

Habitat

Only found near populations of the two parents N. ampullaria and N. rafflesiana.A

Hybrid parent formula

N. ampullaria × N. rafflesiana.C

Bibliography

A. Cheek, M.R. & Jebb, M.H.P. 2001: Flora Malesiana - Nepenthaceae, Series I, Volume 15
B. Berendsohn, W.G. 2017+: Compilation of Nepenthes data for the Caryophyllales taxonomic backbone.
C. Macfarlane, J.M. 1908: pp. 1-92. – In: Engler, A., Pflanzenreich IV. 111 (Heft 36)