Nepenthes villosa

Synonymy

Nepenthes villosa Hook.f. in Hooker, Icon. Pl. 9: t. 888. 1852 sec. Cheek & Jebb 2001

Holotype (designated by Hooker, J.D. 18521): Malaysia, Borneo, Sabah, Mt Kinabalu, alt. 7000-8000 feet, H. Low s.n. (K)

1. Hooker, J.D. 1852 – In: Hooker, W. J., Icones plantarum 9

Content

Other sources

p.p., excl. Nepenthes edwardsiana (Danser, B. H. 1928: – Bull. Jard. Bot. Buitenzorg, sér. 3, 9: 396)

p.p., excl. Nepenthes edwardsiana (Beck von Mannagetta und Lerchenau, G. 1895: Die Gattung Nepenthes. – Wiener Ill. Gart.-Zeitung 20: 96-107, 141-150, 182-192, 217-229: 183)

Adam, J. H. & Wilcock, C.C. 1998: – Sarawak Mus. J. 50: 75-77: 159, f. XXIIId.

Kurata, S. 1976: Nepenthes of Mt Kinabalu, Sabah. – Sabah National Parks Trustees, Kota Kinabalu: 73, t. 25 & 26

Phillipps, A. & Lamb, A.L. 1996: Pitcher Plants of Borneo: 149, f. 79

Jebb, M. H. P. & Cheek, M. R. 1997: A skeletal revision of Nepenthes. – Blumea 42(1): 1-106: 91

Clarke, C. M. 1997: Nepenthes of Borneo. – Kota Kinabalu: Natural History Publications: 135, f. 93 & 94

Description

Terrestrial shrub, not known to climb, 0.6(-1.5) m tall. Stem terete, 6-10 mm diam., often at length prostrate. Leaves coriaceous, petiolate; blade elliptic or oblong, 10-20 by 5-10 cm, apex emarginate, base obtuse; petiole canaliculate, 4-12 cm long, clasping the stem for up to 4/5 its circumference, sheathing, not auriculate or decurrent. Longitudinal nerves 1-3 on each side of the midrib in the outer third, conspicuous. Pennate nerves numerous, arising at 45° from the midrib, reaching the outer longitudinal nerve, conspicuous. Lower pitchers broadly ellipsoidal, 5-16 by 5-12 cm with two fringed wings in the upper 1/3 or 1/2, the wings 5-12 mm wide, fringed elements 2.5-8 mm long, 1-4 mm apart; mouth subcircular, almost horizontal, then abruptly rising to the vertical in the rear to provide a stout column 2-3.5 cm high; peristome rounded, 6-12 mm broad, ribs 3-10 mm apart and 4-6 mm high, outer surface entire, never sinuate, inner surface with teeth 7-12 mm long, in two distinct ranks, strikingly protruding up the column; lid suborbicular 5-12 by 5-12 cm, apex rounded, base conspicuously cordate, lower surface lacking appendages, glands very dense, circular, 0.1-0.3 mm diam.; spur stout, entire, c. 10 mm long. Upper pitchers rarely seen, as the lower pitchers, but ovoid to infundibulate, 10-18 by 5-12 cm. Male inflorescence 35-70 cm long; peduncle 20-50 cm long; partial peduncles 1-flowered, c. 100; pedicels 9-15 mm long; bracts absent; tepals elliptic, 5-7 by 2-3 mm; androphore 2-2.5 mm long; anther head 1-1.5 by 1.5-2 mm. Fruit not seen. Seed not seen. Indumentum villose, hairs simple, 3-4 mm long, red-brown, densely covering all parts of the plant, persistent on lower leaf and leaf edge, tendril and pitcher. Colour of outer pitcher and lid yellow, suffused with pink or red, inner white or pale green, indumentum brown.A

Notes

Nepenthes villosa is closely related and sometimes confused with N. edwardsiana (which also occurs on Mt Kinabalu) and N. macrophylla. It overlaps slightly in altitudinal range with N. edwardsiana but is found in a different habitat (Phillipps & A.L. Lamb, Pitcher Plants of Borneo (1996) 149). Nepenthes edwardsiana is easily distinguished by its more elongated, less hairy pitchers which are slightly constricted about the middle. Intermediate taxa can be ascribed to the hybrid N.harryana Burb. Nepenthes villosa differs from both N. edwardsiana and N. macrophylla in its emarginate leaf blade, long villose tendril, and in its ellipsoid, villose pitcher which lacks any narrowing at its middle. The recently described N. mira of Palawan also seems to be a part of the N. villosa group.
Nepenthes edwardsiana and N. villosa were united by Danser Bull. Jard. Bot. Buitenzorg III 9 (1928). Macfarlane reinstated them in Das Pflanzenreich 4, 3 (1908).A

Hybrids

1. Nepenthes villosa x N. edwardsiana = Nepenthes x harryana (Burb.) Macfarl. (see under Nepenthes edwardsiana).A 2. Nepenthes villosa x N. rajah = Nepenthes x kinabaluensis Sh. Kurata (see Nepenthes x kinabaluensis).A

Distribution (General)

Borneo: Sabah (Mt Kinabalu and Mt Tambuyukon).A

Habitat

Mossy forest with Dacrydium and Leptospermum, or amongst boulders, shrubs and grass; ultramafic substrates; 2400-3200 m.A

Bibliography

A. Cheek, M. R. & Jebb, M. H. P. 2001: Flora Malesiana - Nepenthaceae, Series I, Volume 15. – Leiden: Nationaal Herbarium Nederland, Universiteit Leiden branch

Specimens

	Country	Date	Collector + collecting number	Herbaria	Type	Scan	Derivatives
	Malaysia		Low, H. s.n.	K
	Citation: Malaysia, Borneo, Sabah, Mt Kinabalu, alt. 7000-8000 feet, H. Low s.n. Specimen summary: K Holotype of Nepenthes villosa Hook.f.