Nepenthes naga

Nepenthes naga

Synonymy

Nepenthes naga Akhriadi, Hernawati, Primaldhi & M.Hambali in Reinwardtia 12(5): 340 (-342; fig. 1). 2009 sec. Akhriadi & al. 2009
    • Holotype (designated by Akhriadi, P., Hernawati, Primaldhi, A. & Hambali, M. 20091): Indonesia, North Sumatra, the hill around Panyabungan city, alt. 1500 – 2000 m, 27 Jul 2007, A. Primaldhi & M. Hambali DivNep 052 (ANDA).
  • 1. Akhriadi, P., Hernawati, Primaldhi, A. & Hambali, M. 2009: Nepenthes naga, a new species of Nepenthaceae from Bukit Barisan of Sumatra. – Reinwardtia 12(5): 339-342

Description

Epiphytic climber to ca. 5 m tall. Stem of rosette and lower parts cylindrical, ca. 1 cm in diameter, internodes ca. 0.7 cm length. Stem of upper parts climbing 100–200 cm, orbicular – quadrangular, ca. 0.8 cm in diameter, internodes 6.8–14.8 cm length, with a spine-like process above each nodes. Leaves of rosette and lower parts leathery coriaceous, sessile, spathulate – oblong, 21–27 by 6.0–7.8 cm, base decurrent for 2/3 of its diameter; midrib sunken above, triangular beneath; longitudinal veins 3 pairs at each side of the midrib, indistinct above and distinct beneath; pinnate veins indistinct at both surfaces, margin entire, apex slightly emarginated – rotundate; tendril inserted ca. 0.3 cm below the apex, 18.5–41.0 cm length. Leaves of upper parts similar to those of the rosette and the lower parts; but spathulate, 10.2–16.0 by 4.5–6.0 cm, decurrent along the stem for 1/2–3/4 of its diameter at the base, apex rotundate; tendril insertion apical, looped, 24.0–28.0 cm length. Rosette and lower pitchers ovoid to narrowly ovoid at the base to glandular zone extended 1/2–3/4 pitchers high then narrowly cylindrical towards the mouth, 24.0–33.5 by 5.7–6.8 cm; two wings extended down from the edge of the mouth, 10.5–17.3 by 0.5 cm width, with fringed hairs 1.4–1.9 cm in length; mouth ovate, 6.7–8.4 by 2.4–3.7 cm, slightly necked, 45–60 slope; peristome expanded outwards 2.3–5.8 cm wide at each side, curved downward inside the pitcher 0.4–0.8 cm width, 0.5–0.8 cm width in front, with 6–9 lobes on each side, with 3 notches in front with 0.2–0.5 cm width; teeth distinct 0.05–0.1 cm length, 0.3–0.4 cm length at the neck; lid ovate, 7.3–8.5 by 5.0–7.2 cm, base cordate, midrib sunken–grooved above, raised beneath, longitudinal veins 4 or 5 pairs that insertion from the base, distinct above, indistinct beneath; margin undulate, apex rotun- date; 2 appendages along the midrib of the lid beneath, first appendage sharp like teeth 0.6–1.0 cm distance from the midrib base with 0.4–0.7 cm height; second appendage triangular dichotomous appendages like snake-tongue 1.2–2.3 cm distance inserted apically reach 1.0–1.4 by 0.3–0.5 cm, basal of dichotomous appendages 1.4–2.3 cm width; concentrated nectar gland on lid beneath surface along the midrib 0.01–0.05 cm in diameter, larger nectar gland concentrated at dichotomous appendages 0.05–0.1 cm in diameter; spur 0.3–0.6 cm distance below the lid base, 2.1–2.8 cm length, unbranched. Upper pitchers slightly infundibular at the base to glandular zone 1/3–3/4 pitcher high then cylindrical towards the mouth, 20.8–24.3 by 4.0–4.5 cm; wings reduced to rib; mouth ovate, 3.8–5.4 by 3.0–3.5 cm, necked, 45° slope; peristome expanded outwards 1.0–0.5 cm width on both sides, 0.2–0.3 cm width in front; 4–5 lobes each sides, 0–1 notched in front 0–0.2 cm length; teeth distinct 0.05 cm height, 0.1–0.13 cm height at the neck; lid ovate, 5.5–6.1 cm length, 4.3–5.5 cm width, base cordate; midrib grooved above, raised beneath; longitudinal veins 3–4 pairs, distinct above, indistinct beneath; margin undulate, apex rotundate; 2 appendage along the midrib of the lid beneath, first appendage sharp like teeth 0.5–0.6 cm distance from the midrib base with 0.1–0.3 cm height; second appendage triangular dichotomous like snake-tongue 1.1–1.3 cm distance inserted api- cally, each 1.3–1.5 cm length and 0.3–0.5 cm width each blade, basal of dichotomous appendages 1.4–1.5 cm width; concentrated nectar gland on lid beneath surface along the midrib 0.01–0.02 cm in diameter, bigger nectar gland concentrated at dichotomous appendages surface 0.05–0.1 cm in diameter; spur flattened, 1.0–1.4 cm length, 0.2 cm width, 0.4–0.6 cm distance below the lid base, 2 branched near the tip. Female flowers axillary in front of the leaf base, a raceme or a panicle, 14.5 cm length, peduncle 7 cm length, rachis 7.5 cm length, pedicel 0.3–0.5 cm length, pedicel branches 0.5–0.9 cm length; bracteole linear, 0.7–1.2 cm length; tepal linear 0.2–0.5 cm length. Fruit 0.3–1.0 cm length, 0.15–0.4 cm width. Male flowers not found. Indumen- tum of the rosette and lower parts glabrous. Indumentum of the upper parts similar to those of the rosette and lower parts including the female flowers. Colour of herbarium specimen stem blackish brown, leaves above greenish brown and brown beneath, pitchers dark brown in rosette and lower pitcher and bright brown in upper pitcher, lid greenish brown – brown above and dark brown beneath. Colour of living specimen leaves dark green above and pale green beneath, midrib pale green on both sides. Rosette and lower pitchers green – brownish green outside, glandular zone pale green inside, upper zone pale green with dark red blotches inside, mouth dark red – blackish brown; lid green with dark red blotches around the midrib then dark red spots to the margin above, green with dark blotches beneath, teeth-like appendage green on the base of lid beneath, dichotomous appendage dark red on the apical below of lid beneath. Upper pitchers similar to those of the rosette and the lower pitchers, but green at glandular zone outside and with pale dark red toward the mouth, mouth green with dark red lines, lid green – green with dark red spots above, pale green with dark red blotches beneath, dichotomous appendage green with dark red blotches. Fruit brown.A

Distribution (General)

Indonesia, Sumatra, North SumatraA

Etymology

The specific epithet naga refers to the dichotomous appendage like snake's tongue inserted sub-apically on the under surface of the lid. Local folklore also includes stories claiming that the habitat of this species was occupied by dragons (‘naga’), long ago.A

Habitat

Epiphytic in montane mossy forest at 1,500 to 2,000 m a.s.l.A

Bibliography

A. Akhriadi, P., Hernawati, Primaldhi, A. & Hambali, M. 2009: Nepenthes naga, a new species of Nepenthaceae from Bukit Barisan of Sumatra. – Reinwardtia 12(5): 339-342

Specimens

CountryDateCollector + collecting numberHerbariaTypeScanDerivatives
Indonesia2007-07-27Primaldhi, A. & Hambali, M. DivNep 052ANDA
Citation: Indonesia, North Sumatra, the hill around Panyabungan city, alt. 1500 – 2000 m, 27 Jul 2007, A. Primaldhi & M. Hambali DivNep 052.

Specimen summary: ANDA
Holotype of Nepenthes naga Akhriadi, Hernawati, Primaldhi & M.Hambali