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Achatocarpaceae Heimerl, Nat. Pflanzenfam. ed. 2. 16c: 174. 1934, nom. cons., sec. APG IV [Angiosperm Phylogeny Group] 20161
  • 1. APG IV [Angiosperm Phylogeny Group] 2016: APG IV: An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants. – Botanical Journal of the Linnean Society 181(1): 1-20.


A small family comprising two genera and 16 species occurring in tropical America, from southeastern United States to South America (Medina 2009). Traditionally, the family has been included in Phytolaccaceae s.l., but its position as an independent lineage has been well-supported by several molecular phylogenetic studies (Hilu & al. 2003; Schäferhoff & al. 2009; Brockington & al. 2009, 2011), which also showed that the family is more closely related to the Amaranthaceae/Chenopodiaceae clade rather than to Phytolaccaceae. Achatocarpaceae are characterised by having unisexual flowers, the gynoecium with two connate carpels, unilocular ovaries with two styles and a single ovule, berrylike fruits and pollen with obscure pores (Martínez-García 1985; Lipscomb 2003).A,B,C,D,E,F,G,H


A. Brockington, S. F., Alexandre, R., Ramdial, J., Moore, M. J., Crawley, S., Dhingra, A., Hilu, K., Douglas, E. S. & Soltis, P.S. 2009: Phylogeny of the Caryophyllales sensu lato: Revisiting hypotheses on pollination biology and perianth differentiation in the core Caryophyllales. – International Journal of Plant Sciences 170(5): 627–643.
B. Brockington, S. F., Walker, R. H., Glover, R. H., Soltis, P.S. & Soltis, D.E. 2011: Complex pigment evolution in the Caryophyllales. – New Phytologist 190: 854 – 864.|
C. Hernández-Ledesma, P., Berendsohn, W. G., Borsch, T., von Mering, S., Akhani, H., Arias, S., Castañeda-Noa, I., Eggli, U., Eriksson, R., Flores-Olvera, H., Fuentes-Bazan, S., Kadereit, G., Klak, C., Korotkova, N., Nyffeler, R., Ocampo, G. & Ochoterena, H. 2015: A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales. – Willdenowia 45(3): 281-383.
D. Hilu, K., Müller, K. F., Soltis, D. E., Soltis, P. S., Savolainen, V., Chase, M. W., Powell, M. P., Lawrence, Alice, Evans, R. C., Sauquet, H., Neinhuis, C., Slotta, T., Rohwer, J. G., Campbell, C. S. & Chatrou, L. 2003: Angiosperm phylogeny based on matK sequence information. – American Journal of Botany 90: 1758 – 1776.
E. Lipscomb, B. L. 2004: Achatocarpaceae. Flora of North America 4: 13-14
F. Martínez-García, J. 1985: Achatocarpaceae, 1 – 9. – In: Gómez-Pompa, A. (ed.), Flora de Veracruz 45. – Veracruz: Instituto Nacional de Investigaciones sobre Recursos Bióticos
G. Medina, L. R. 2009: Achatocarpaceae, 1 – 5. – In: Flora del Valle de Tehuacán Cuicatlán 73. – México D. F.: Instituto de Biología, Universidad Nacional Autónoma de México
H. Schäferhoff, B., Müller, K.F. & Borsch, T. 2010: Caryophyllales phylogenetics: disentangling Phytolaccaceae and Molluginaceae and description of Microteaceae as a new isolated family. – Willdenowia 39: 209-228.