Philippines, Mindanao Island, Davao Oriental Province, Mount Hamiguitan, trail from San Isidro in mountain forest margins on the summit ridge, 1310 m, March 13, 2007. V. Amoroso and R. Aspiras CMUH00006494, Central Mindanao Univierstiy Herbarium, Musuan, Bukidnon, Philippines (CMUH).