Record basis:

Preserved Specimen

Specimen type designations:

• Neotype (designated by Sukhorukov, A. P., Sennikov, A.N., Nilova, M.V., Mazei, Y., Kushunina, M., Marchioretto, M.S. & Hanáček, P. 2019: p. 24 (designated by Sennikov & Sukhorukov)1)